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IAWA List of Microscopic Features for Softwood Identification (eds. H.G. Richter, D. Grosser, I. Heinz, P.E. Gasson)
IAWA J. 25 (1): 1-70
[Also separately obtainable from the IAWA Office in Leiden, Netherlands, for the price of EUR 25.00 or USD 30.00, excluding postage: Click here for the order form]
Preface A definitive list of anatomical features of softwoods has long been needed. The hardwood list (IAWA Committee 1989) has been adopted throughout the world, not least because it provides a succinct, unambiguous illustrated glossary of hardwood characters that can be used for a variety of purposes, not just identification. This publication is intended to do the same job for softwoods. Identifying softwoods relies on careful observation of a number of subtle characters, and great care has been taken to show high quality photomicrographs that remove most of the ambiguity that definitions alone would provide.
Unlike the Hardwood Committee, the Softwood Committee never met in its full composition. The softwood committee members attending the XVI International Botanical Congress in St. Louis in August 1999 met for a day to discuss an early draft. The editing of the list was coordinated by Jorgo (H.G.) Richter, who intensively communicated by e-mail with all committee members. At the IAWA meeting in Portland, Oregon in July 2003, several committee members discussed a late draft. All the photographs were taken by Dietger Grosser and Immo Heinz, whose MSc thesis work was the catalyst for the formation of the committee. Peter Gasson then did the final editing and cross-referencing of text and plates before sending the entire work to Leiden for publication.
We hope that all present and future colleagues engaged in wood identification and descriptive wood anatomy will find this list a valuable guide and reference.
LIST OF FEATURES
Name (nomenclature)
GENERAL INFORMATION
Geographical distribution p. 12
1. Europe and temperate Asia (Brazier and Franklin region 74)
2. Europe, excl. Mediterranean
3. Mediterranean incl. Northern Africa and Middle East
4. Temperate Asia (China, Japan, Russia)
5. Central South Asia (Brazier and Franklin region 75)
6. India, Pakistan, Sri Lanka
7. Burma (Myanmar)
8. Southeast Asia and the Pacific (Brazier and Franklin region 76)
9. Thailand, Laos, Vietnam, Cambodia (Indochina)
10. Indomalesia (Indonesia, Philippines, Malaysia, Brunei, Singapore, Papua New Guinea, and Solomon Islands)
11. Pacific Islands (incl. New Caledonia, Samoa, Hawaii, and Fiji)
12. Australia and New Zealand (Brazier and Franklin region 77)
13. Australia
14. New Zealand
15. Tropical mainland Africa and adjacent islands (Brazier and Franklin region 78)
16. Tropical Africa
17. Madagascar, Mauritius, Reunion and Comores
18. Southern Africa (south of the Tropic of Capricorn) (Brazier and Franklin region 79)
19. North America (north of Mexico) (Brazier and Franklin region 80)
20. Neotropics and temperate Brazil (Brazier and Franklin region 81)
21. Mexico and Central America
22. Caribbean
23. Tropical South America
24. Southern Brazil
25. Temperate South America including Argentina, Chile, Uruguay and southern Paraguay (Brazier and Franklin region 82)
PHYSICAL PROPERTIES
Heartwood colour p. 12
26. Brown or shades of brown
27. Red or shades of red
28. Yellow or shades of yellow
29. Light coloured (whitish, creamy, grey)
30. Purple or shades of purple
31. Other than above (specify)
Difference between heartwood and sapwood colour p. 14
32. Heartwood colour similar to sapwood colour
33. Heartwood colour distinct from sapwood colour
Presence of heartwood with colour streaks p. 14
34. Heartwood with streaks
Presence of a distinct odour p. 14
35. Odour distinct (specify)
Average air-dry density / basic specific gravity p. 15
36.
< g/cm3 >
Average air-dry density [g/cm3] (categories) p. 15
37. Less than 0.48 g/cm3
38. 0.480.60 g/cm3
39. Above 0.60 g/cm3
GROWTH RINGS
Presence of growth ring boundaries p. 16
40. Growth ring boundaries distinct
41. Growth ring boundaries indistinct or absent
Transition from earlywood to latewood p. 16
42. Abrupt
43. Gradual
TRACHEIDS
Tracheid pitting in radial walls (in earlywood only) p. 19
44. (predominantly) Uniseriate
45. (predominantly) Two or more seriate
Arrangement of (two or more seriate) tracheid pitting in radial walls (earlywood only) p. 19
46. Opposite
47. Alternate
Organic deposits (in heartwood tracheids) p. 21
48. Present
Average tracheid length p. 22 49.
< µm >
Average tracheid length (size classes) p. 22 50. Short (less than 3000 µm) 51. Medium (3000 to 5000 µm) 52. Long (over 5000 µm)
Intercellular spaces throughout the wood (in transverse section) p. 23
53. Present
Latewood tracheid wall thickness p. 24
54. Thin-walled (double wall thickness less than radial lumen diameter)
55. Thick-walled (double wall thickness larger than radial lumen diameter)
Torus (pits in earlywood tracheids only) p. 25
56. Present
57. Scalloped
Torus extensions p. 26
58. Present
Pits with notched borders p. 28
59. Present
Warty layer (visible under the light microscope) p. 28
60. Present
HELICAL AND OTHER WALL THICKENINGS
Helical thickenings in tracheids p. 30
Helical thickenings in longitudinal tracheids (presence) p. 32
61. Present
Helical thickenings (in longitudinal tracheids - location) p. 32
62. Present throughout the growth increment
63. Well developed only in earlywood
64. Well developed only in latewood
Helical thickenings (in longitudinal tracheids - whether single or grouped)
p. 32
65. Single
66. Grouped (double or triple)
Helical thickenings (in longitudinal tracheids - spacing, earlywood tracheids only) p. 32 67. Narrowly spaced (number of coils more than 120 per mm) 68. Widely spaced (number of coils less than 120 per mm)
Helical thickenings in ray tracheids p. 33
69. Commonly present
70. (present but) Rare
Callitroid thickenings p. 34
71. Present
AXIAL PARENCHYMA
Axial parenchyma (excl. epithelial and subsidiary cells of intercellular canals) p. 35
72. Present
Arrangement of axial parenchyma p. 37
73. Diffuse (evenly scattered throughout the entire growth increment)
74. Tangentially zonate
75. Marginal
Transverse end walls p. 39
76. Smooth
77. Irregularly thickened
78. Beaded or nodular
RAY COMPOSITION
Ray tracheids p. 40
79. Commonly present
80. Absent or very rare
Cell walls of ray tracheids p. 43
81. Smooth
82. Dentate
83. Reticulate
Ray tracheid pit borders angular or with dentate thickenings (radial section, Larix & Picea only) p. 45
84. Present
End walls of ray parenchyma cells p. 47
85. Smooth (unpitted)
86. Distinctly pitted
Horizontal walls of ray parenchyma cells p. 48
87. Smooth (unpitted)
88. Distinctly pitted
Indentures p. 49
89. Present
CROSS-FIELD PITTING
Cross-field pitting (according to Phillips 1948, amended by Vogel 1995) p. 51
90. Window-like (fenestriform)
91. Pinoid
92. Piceoid
93. Cupressoid
94. Taxodioid
95. Araucarioid
Number of pits per cross-field (earlywood tracheids only) p. 54
96.
< number per crossfield >
Number of pits per cross-field (earlywood only categories) p. 54
97. (large window-like) 12
98. 13
99. 35
100. 6 or more
RAY SIZE
Average ray height p. 55 101.
< µm >
Average ray height (number of cells) p. 57
102. Very low (up to 4 cells)
103. Medium (5 to 15 cells)
104. High (from 16 to 30 cells)
105. Very high (more than 30 cells)
Average fusiform ray height p. 57 106.
< µm >
Ray width (cells) p. 57
107. Exclusively uniseriate
108. 23-seriate in part
INTERCELLULAR CANALS
Axial intercellular (resin) canals p. 58
109. Present
Radial intercellular (resin) canals p. 60
110. Present
Traumatic (resin) canals (axial, radial) p. 60
111. Present
Average diameter of normal axial intercellular canals p. 60 112. Tangential diameter, delimited by epithelial cells (Method A) < µm > 113. Tangential diameter of entire resin canal complex (Method B) < µm > 114. Radial diameter, delimited by epithelial cells (Method C) < µm >
Average diameter of normal radial intercellular canals p. 62 115.
< µm >
Epithelial cells (of intercellular canals) p. 62
116. Thick-walled
117. Thin-walled
MINERAL INCLUSIONS
Crystals p. 65
118. Present
Type of crystals p. 65
119. Prismatic
120. Druses
121. Other forms (specify)
Crystals located in p. 65
122. Rays
123. Axial parenchyma
124. Cells associated with intercellular canals
The IAWA Committee:
PIETER BAAS
Nationaal Herberium Nederland, Universiteit Leiden branch, The Netherlands
baas@nhn.leidenuniv.nl
NADEZHDA BLOKHINA
Institute of Biology and Pedology, Far East branch, Russian Academy of Science, Vladivostok, Russia
evolut@eastnet.febras.ru
TOMOYUKI FUJII
Forestry & Forest Products Research Institute, Ibaraki, Japan
tfujii@ffpri.affrc.go.jp
PETER E. GASSON
Jodrell Laboratory, Royal Botanic Gardens, Kew, U.K.
p.gasson@kew.org
DIETGER GROSSER
Institut für Holzforschung der Universität München, Germany
grosser@holz.forst.uni-muenchen.de
IMMO HEINZ
Institut für Holzforschung der Universität München, Germany
heinz@holz.forst.tu-muenchen.de
JUGO ILIC
CSIRO Forestry & Forest Products, South Clayton, Australia
jugo.ilic@ffp.csiro.au
JIANG XIAOMEI
Chinese Research Institute of Wood Industry (CRIWI), Chinese Academy of Forestry, Beijing, China
xiaomei@wood.forestry.ac.cn
REGIS B. MILLER
USDA Forest Service, Forest Products Laboratory, Madison, Wisconsin, U.S.A.
rmiller1@wisc.edu
LEE ANN NEWSOM
Department of Anthropology, Pennsylvania State University, U.S.A.
lan12@psu.edu
SHUICHI NOSHIRO
Forestry & Forest Products Research Institute, Ibaraki, Japan
noshiro@ffpri.affrc.go.jp
HANS GEORG RICHTER
Institut für Holzbiologie, Universität Hamburg, Germany
hrichter@holz.uni-hamburg.de
MITSUO SUZUKI
Botanical Garden, Graduate School of Science, Tohoku University, Sendai, Japan
mitsuos@mail.cc.tohoku.ac.jp
TERESA TERRAZAS
Colegio de Postgraduados, Programa de Botánica, Montecillo, Mexico
winchi@colpos.colpos.mx
ELISABETH A. WHEELER
Department of Wood & Paper Science, North Carolina State University, Raleigh, North Carolina, U.S.A.
xylem@unity.ncsu.edu
ALEX C. WIEDENHOEFT
USDA Forest Service, Forest Products Laboratory, Madison, Wisconsin, U.S.A.
acwieden@wisc.edu
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