Abstracts IAWA Journal 25 (1)

IAWA List of Microscopic Features for Softwood Identification (eds. H.G. Richter, D. Grosser, I. Heinz, P.E. Gasson)
IAWA J. 25 (1): 1-70
[Also separately obtainable from the IAWA Office in Leiden, Netherlands, for the price of EUR 25.00 or USD 30.00, excluding postage: Click here for the order form]

Preface – A definitive list of anatomical features of softwoods has long been needed. The hardwood list (IAWA Committee 1989) has been adopted throughout the world, not least because it provides a succinct, unambiguous illustrated glossary of hardwood characters that can be used for a variety of purposes, not just identification. This publication is intended to do the same job for softwoods. Identifying softwoods relies on careful observation of a number of subtle characters, and great care has been taken to show high quality photomicrographs that remove most of the ambiguity that definitions alone would provide.
Unlike the Hardwood Committee, the Softwood Committee never met in its full composition. The softwood committee members attending the XVI International Botanical Congress in St. Louis in August 1999 met for a day to discuss an early draft. The editing of the list was coordinated by Jorgo (H.G.) Richter, who intensively communicated by e-mail with all committee members. At the IAWA meeting in Portland, Oregon in July 2003, several committee members discussed a late draft. All the photographs were taken by Dietger Grosser and Immo Heinz, whose MSc thesis work was the catalyst for the formation of the committee. Peter Gasson then did the final editing and cross-referencing of text and plates before sending the entire work to Leiden for publication.
We hope that all present and future colleagues engaged in wood identification and descriptive wood anatomy will find this list a valuable guide and reference.


LIST OF FEATURES

Name (nomenclature)

GENERAL INFORMATION

Geographical distribution — p. 12
1. Europe and temperate Asia (Brazier and Franklin region 74)
2. Europe, excl. Mediterranean
3. Mediterranean incl. Northern Africa and Middle East
4. Temperate Asia (China, Japan, Russia)
5. Central South Asia (Brazier and Franklin region 75)
6. India, Pakistan, Sri Lanka
7. Burma (Myanmar)
8. Southeast Asia and the Pacific (Brazier and Franklin region 76)
9. Thailand, Laos, Vietnam, Cambodia (Indochina)
10. Indomalesia (Indonesia, Philippines, Malaysia, Brunei, Singapore, Papua New Guinea, and Solomon Islands)
11. Pacific Islands (incl. New Caledonia, Samoa, Hawaii, and Fiji)
12. Australia and New Zealand (Brazier and Franklin region 77)
13. Australia
14. New Zealand
15. Tropical mainland Africa and adjacent islands (Brazier and Franklin region 78)
16. Tropical Africa
17. Madagascar, Mauritius, Reunion and Comores
18. Southern Africa (south of the Tropic of Capricorn) (Brazier and Franklin region 79)
19. North America (north of Mexico) (Brazier and Franklin region 80)
20. Neotropics and temperate Brazil (Brazier and Franklin region 81)
21. Mexico and Central America
22. Caribbean
23. Tropical South America
24. Southern Brazil
25. Temperate South America including Argentina, Chile, Uruguay and southern Paraguay (Brazier and Franklin region 82)

PHYSICAL PROPERTIES

Heartwood colour — p. 12
26. Brown or shades of brown
27. Red or shades of red
28. Yellow or shades of yellow
29. Light coloured (whitish, creamy, grey)
30. Purple or shades of purple
31. Other than above (specify)

Difference between heartwood and sapwood colour — p. 14
32. Heartwood colour similar to sapwood colour
33. Heartwood colour distinct from sapwood colour

Presence of heartwood with colour streaks — p. 14
34. Heartwood with streaks

Presence of a distinct odour — p. 14
35. Odour distinct (specify)

Average air-dry density / basic specific gravity — p. 15
36. … < g/cm3 >

Average air-dry density [g/cm3] (categories) — p. 15
37. Less than 0.48 g/cm3
38. 0.48–0.60 g/cm3
39. Above 0.60 g/cm3

GROWTH RINGS

Presence of growth ring boundaries — p. 16
40. Growth ring boundaries distinct
41. Growth ring boundaries indistinct or absent

Transition from earlywood to latewood — p. 16
42. Abrupt
43. Gradual

TRACHEIDS

Tracheid pitting in radial walls (in earlywood only) — p. 19
44. (predominantly) Uniseriate
45. (predominantly) Two or more seriate

Arrangement of (two or more seriate) tracheid pitting in radial walls (earlywood only) — p. 19
46. Opposite
47. Alternate

Organic deposits (in heartwood tracheids) — p. 21
48. Present

Average tracheid length — p. 22
49. … < µm >

Average tracheid length (size classes) — p. 22
50. Short (less than 3000 µm)
51. Medium (3000 to 5000 µm)
52. Long (over 5000 µm)

Intercellular spaces throughout the wood (in transverse section) — p. 23
53. Present

Latewood tracheid wall thickness — p. 24
54. Thin-walled (double wall thickness less than radial lumen diameter)
55. Thick-walled (double wall thickness larger than radial lumen diameter)

Torus (pits in earlywood tracheids only) — p. 25
56. Present
57. Scalloped

Torus extensions — p. 26
58. Present

Pits with notched borders — p. 28
59. Present

Warty layer (visible under the light microscope) — p. 28
60. Present

HELICAL AND OTHER WALL THICKENINGS

Helical thickenings in tracheids — p. 30

Helical thickenings in longitudinal tracheids (presence) — p. 32
61. Present

Helical thickenings (in longitudinal tracheids - location) — p. 32
62. Present throughout the growth increment
63. Well developed only in earlywood
64. Well developed only in latewood

Helical thickenings (in longitudinal tracheids - whether single or grouped) —
p. 32
65. Single
66. Grouped (double or triple)

Helical thickenings (in longitudinal tracheids - spacing, earlywood tracheids only) — p. 32
67. Narrowly spaced (number of coils more than 120 per mm)
68. Widely spaced (number of coils less than 120 per mm)

Helical thickenings in ray tracheids — p. 33
69. Commonly present
70. (present but) Rare

Callitroid thickenings — p. 34
71. Present

AXIAL PARENCHYMA

Axial parenchyma (excl. epithelial and subsidiary cells of intercellular canals) — p. 35
72. Present

Arrangement of axial parenchyma — p. 37
73. Diffuse (evenly scattered throughout the entire growth increment)
74. Tangentially zonate
75. Marginal

Transverse end walls — p. 39
76. Smooth
77. Irregularly thickened
78. Beaded or nodular

RAY COMPOSITION

Ray tracheids — p. 40
79. Commonly present
80. Absent or very rare

Cell walls of ray tracheids — p. 43
81. Smooth
82. Dentate
83. Reticulate

Ray tracheid pit borders angular or with dentate thickenings (radial section, Larix & Picea only) — p. 45
84. Present

End walls of ray parenchyma cells — p. 47
85. Smooth (unpitted)
86. Distinctly pitted

Horizontal walls of ray parenchyma cells — p. 48
87. Smooth (unpitted)
88. Distinctly pitted

Indentures — p. 49
89. Present

CROSS-FIELD PITTING

Cross-field pitting (according to Phillips 1948, amended by Vogel 1995) — p. 51
90. “Window-like” (fenestriform)
91. Pinoid
92. Piceoid
93. Cupressoid
94. Taxodioid
95. Araucarioid

Number of pits per cross-field (earlywood tracheids only) — p. 54
96. … < number per crossfield >

Number of pits per cross-field (earlywood only – categories) — p. 54
97. (large window-like) 1–2
98. 1–3
99. 3–5
100. 6 or more

RAY SIZE

Average ray height — p. 55
101. … < µm >

Average ray height (number of cells) — p. 57
102. Very low (up to 4 cells)
103. Medium (5 to 15 cells)
104. High (from 16 to 30 cells)
105. Very high (more than 30 cells)

Average fusiform ray height — p. 57
106. … < µm >

Ray width (cells) — p. 57
107. Exclusively uniseriate
108. 2–3-seriate in part

INTERCELLULAR CANALS

Axial intercellular (resin) canals — p. 58
109. Present

Radial intercellular (resin) canals — p. 60
110. Present

Traumatic (resin) canals (axial, radial) — p. 60
111. Present

Average diameter of normal axial intercellular canals — p. 60
112. Tangential diameter, delimited by epithelial cells (Method A) < µm >
113. Tangential diameter of entire resin canal complex (Method B) < µm >
114. Radial diameter, delimited by epithelial cells (Method C) < µm >

Average diameter of normal radial intercellular canals — p. 62
115. … < µm >

Epithelial cells (of intercellular canals) — p. 62
116. Thick-walled
117. Thin-walled

MINERAL INCLUSIONS

Crystals — p. 65
118. Present

Type of crystals — p. 65
119. Prismatic
120. Druses
121. Other forms (specify)

Crystals located in — p. 65
122. Rays
123. Axial parenchyma
124. Cells associated with intercellular canals



The IAWA Committee:
PIETER BAAS
Nationaal Herberium Nederland, Universiteit Leiden branch, The Netherlands
baas@nhn.leidenuniv.nl

NADEZHDA BLOKHINA
Institute of Biology and Pedology, Far East branch, Russian Academy of Science, Vladivostok, Russia
evolut@eastnet.febras.ru

TOMOYUKI FUJII
Forestry & Forest Products Research Institute, Ibaraki, Japan
tfujii@ffpri.affrc.go.jp

PETER E. GASSON
Jodrell Laboratory, Royal Botanic Gardens, Kew, U.K.
p.gasson@kew.org

DIETGER GROSSER
Institut für Holzforschung der Universität München, Germany
grosser@holz.forst.uni-muenchen.de

IMMO HEINZ
Institut für Holzforschung der Universität München, Germany
heinz@holz.forst.tu-muenchen.de

JUGO ILIC
CSIRO Forestry & Forest Products, South Clayton, Australia
jugo.ilic@ffp.csiro.au

JIANG XIAOMEI
Chinese Research Institute of Wood Industry (CRIWI), Chinese Academy of Forestry, Beijing, China
xiaomei@wood.forestry.ac.cn

REGIS B. MILLER
USDA Forest Service, Forest Products Laboratory, Madison, Wisconsin, U.S.A.
rmiller1@wisc.edu

LEE ANN NEWSOM
Department of Anthropology, Pennsylvania State University, U.S.A.
lan12@psu.edu

SHUICHI NOSHIRO
Forestry & Forest Products Research Institute, Ibaraki, Japan
noshiro@ffpri.affrc.go.jp

HANS GEORG RICHTER
Institut für Holzbiologie, Universität Hamburg, Germany
hrichter@holz.uni-hamburg.de

MITSUO SUZUKI
Botanical Garden, Graduate School of Science, Tohoku University, Sendai, Japan
mitsuos@mail.cc.tohoku.ac.jp

TERESA TERRAZAS
Colegio de Postgraduados, Programa de Botánica, Montecillo, Mexico
winchi@colpos.colpos.mx

ELISABETH A. WHEELER
Department of Wood & Paper Science, North Carolina State University, Raleigh, North Carolina, U.S.A.
xylem@unity.ncsu.edu

ALEX C. WIEDENHOEFT
USDA Forest Service, Forest Products Laboratory, Madison, Wisconsin, U.S.A.
acwieden@wisc.edu

 

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Comments on the content: Steven Jansen - Laboratory of Plant Systematics
Production: Steven Jansen, Peter Schols & Steven Dessein | Most recent update: October 10, 2003
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