Research

Publications

Press & media coverage

Talks

Teaching materials

My research centers on social evolution, with particular focus on conflict resolution in social groups. Understanding conflict resolution is considered the key to understanding how stable social groups evolved at all levels of biological organization, including the origin of genomes, multicellular organisms and societies. So far, my main model system has been social insects (ants, bees and wasps). My work has led to various original insights and discoveries, e.g. that cooperation in insect societies is to a large extent socially enforced and that some conflicts in insect societies resemble “tragedies of the commons”, which previously were known mainly from human systems. My research has inspired major new lines of research, and successfully combines a range of techniques including theoretical modelling (game theory, population genetic and inclusive fitness models), video-assisted behavioural observation, chemical ecology (gas chromatography) and molecular techniques (microsatellite genotyping and microarray analyses). Currently I am extending my research towards the study of interspecific mutualisms (using ants and aphids as the main model system, collaboration with W. van den Ende, Plant Physiology, KULeuven), and also into the realm of genomics, to get a grip on what genes are involved in social behaviour and taking advantage of the recently sequenced genome of the honeybee (joint project with Liliane Schoofs and Peter Verleyen, Lab. of Functional Genomics and Proteomics, KULeuven and Dirk de Graaf, UGhent, and in collaboration with the Flemish Institute for Biotechnology MicroArray Facility). Other interests of mine include human social behaviour and experimental evolution studies of social behaviour in microorganisms.

Areas of research: theoretical biology, evolutionary biology, behavioural ecology, sociobiology, sociogenomics, entomology, molecular ecology

52. D.A. Alves, V.L. Imperatriz-Fonseca, T.M. Francoy, P.S. Santos-Filho, P. Nogueira-Neto, J. Billen & T. Wenseleers (2009) The queen is dead – long live the workers: intraspecific parasitism by workers in the stingless bee Melipona scutellaris. Molecular Ecology 18: 4102-4111. See also the News & Views article by Oldroyd & Beekman in the same issue.

51. E. Lommelen, T. Wenseleers, C.A. Johnson, F.P. Drijfhout, J. Billen & B. Gobin (2009) A combination of fertility signals and aggression regulates reproduction in the ant Gnamptogenys striatula. Journal of Insect Behavior, in press.

50. Wenseleers T., Gardner A. & Foster K. R. (2009) Social evolution theory: a review of methods and approaches. In: Social behaviour: genes, ecology and evolution (Tamas Szekely, Allen J. Moore & Jan Komdeur, eds.). Cambridge University Press, 350 pp., ISBN 9780521709620. Order a copy at Amazon.

49. T. Wenseleers (2009) The superorganism revisited. BioScience 59: 702-705. (Review of The Superorganism: The Beauty, Elegance and Strangeness of Insect Societies by B. Hölldobler & E.O. Wilson and The Buzz about Bees: Biology of a Superorganism by J. Tautz)

48. T. Wenseleers (2009) The Superorganism: The Beauty, Elegance and Strangeness of Insect Societies. Quarterly Review of Biology, in press. (Review of B. Hölldobler & E.O. Wilson's new book)

47. T. Wenseleers (2009) De collectieve intelligentie van sociale-insectenkolonies. Karakter 28: 9-11.

46. L. de Meester & T. Wenseleers (2009) Biologische evolutie. In: Wetenschap van het Leven (B. de Groef B. & P. Roels, eds.), pp. 115-139, Acco, Leuven, ISBN 9789033475382. (only intranet access) Order a copy here.

45. P. Roels, T. Wenseleers & B. De Groef (2009) De mens. In: Wetenschap van het Leven (B. de Groef B. & P. Roels, eds.), pp. 177-196, Acco, Leuven, ISBN 9789033475382. (only intranet access) Order a copy here.

44. W. Bonckaert, K. Vuerinckx, J. Billen, R.L. Hammond, L. Keller & T. Wenseleers (2008) Worker policing in the german wasp Vespula germanica. Behavioral Ecology 19: 272-278.

43. M.J. Couvillon, T. Wenseleers, V.L. Imperatriz-Fonseca, P. Nogueira-Neto & F.L.W. Ratnieks (2008) Comparative study in stingless bees (Meliponini) demonstrates that nest entrance size predicts traffic and defensivity. Journal of Evolutionary Biology 21: 194-201.

42. J. Tobback, K. Heylen, B. Gobin, T. Wenseleers, J. Billen, L. Arckens & R. Huybrechts (2008) Cloning and expression of PKG, a candidate foraging regulating gene in Vespula vulgaris. Animal Biology 58: 341-351.

41. F.L.W. Ratnieks & T. Wenseleers (2008) Altruism in insect societies and beyond: voluntary or enforced? Trends in Ecology and Evolution 23: 45-52.

40. T. Wenseleers (2007) Nepotism absent in insect societies – or is it? Molecular Ecology 16: 3063-3065.

39. T. Wenseleers & F.L.W. Ratnieks (2006) Enforced altruism in insect societies. Nature 444: 50. See also the News & Views article in the same issue by D.C. Queller and Nature's Podcast for an interview with coauthor Francis Ratnieks

38. K.R. Foster, T. Wenseleers, F.L.W. Ratnieks & D.C. Queller (2006) There is nothing wrong with inclusive fitness. Trends in Ecology and Evolution 21: 599-560. (reply to letter by Fletcher J.A. et al.)

37. H. Helanterä, A. Tofilski, T. Wenseleers & F.L.W. Ratnieks (2006) Worker policing in the common wasp Vespula vulgaris is not aimed at improving colony hygiene. Insectes Sociaux 53: 399-402.

36. T. Wenseleers & F.L.W. Ratnieks (2006) Comparative analysis of worker reproduction and policing in eusocial Hymenoptera supports relatedness theory. American Naturalist 168: E163-E179.

35. T. Wenseleers (2006) Modelling social evolution: the relative merits and limitations of a Hamilton's rule-based approach. Journal of Evolutionary Biology 19: 1419-1422.

34. P. D’Ettorre, T. Wenseleers, J. Dawson, S. Hutchinson, T. Boswell & F.L.W. Ratnieks (2006) Wax combs mediate nestmate recognition by guard honeybees. Animal Behaviour 71: 773-779.

33. K.R Foster, T. Wenseleers & F.L.W. Ratnieks (2006) Kin selection is the key to altruism. Trends in Ecology and Evolution 21: 57-60. Ranked number 2 in Science Direct's "TOP 25 Hottest Articles"

32. K.R. Foster & T. Wenseleers (2006) A general model for the evolution of mutualisms. Journal of Evolutionary Biology 19: 1283-1293.

31. E. Lommelen, C.A. Johnson, F.P. Drijfhout, J. Billen, T. Wenseleers & B. Gobin (2006) Cuticular hydrocarbons provide reliable cues of fertility in the ant Gnamptogenys striatula. Journal of Chemical Ecology 32: 2023-2034. The original publication is available at www.springerlink.com

30. E.D. Morgan, S.J. Keegans, J. Tits, T. Wenseleers & J. Billen (2006) Preferences and differences in the trail pheromone of the leaf-cutting ant Atta sexdens sexdens (Hymenoptera: Formicidae). European Journal of Entomology 103: 553-558.

29. F.L.W. Ratnieks, K.R. Foster & T. Wenseleers (2006) Conflict resolution in insect societies. Annual Review of Entomology 51: 581-608. (invited review)

28. M.F. Ribeiro, T. Wenseleers, P.S. Santos Filho & D.A. Alves (2006) Miniature queens in stingless bees: basic facts and evolutionary hypotheses. Apidologie 37: 191-206. (invited review, in special issue "Stingless Bees: Biology and Management")

27. E. Schoeters & T. Wenseleers (2005) Onze sociale wespen ("Our social wasps"). Educatie Limburgs Landschap, Heusden Zolder, 135 p. Richly illustrated field guide of the social wasps of Belgium. Price: 10 €. Ordering information

26. T. Wenseleers, N.S. Badcock, K. Erven, A. Tofilski, F.S. Nascimento, A.G. Hart, T.A. Burke, M.E. Archer & F.L.W. Ratnieks (2005) A test of worker policing theory in an advanced eusocial wasp, Vespula rufa. Evolution 59: 1306-1314.
Movie of queen policing in Vespula rufa (mpeg format)
Movie of queen policing in Vespula rufa (windows media format)

25. T. Wenseleers, F.L.W. Ratnieks, M. de F. Ribeiro, D. de A. Alves & V.L. Imperatriz-Fonseca (2005) Working-class royalty: bees beat the caste system. Proceedings of the Royal Society of London - Biology Letters 1: 125-128.

24. T. Wenseleers, A. Tofilski & F.L.W. Ratnieks (2005) Queen and worker policing in the tree wasp Dolichovespula sylvestris. Behavioral Ecology & Sociobiology 58: 80-86.

23. F.L.W. Ratnieks & T. Wenseleers (2005) Policing insect societies. Science 307: 54-56.

22. P. D'Ettorre, E. Brunner, T. Wenseleers & J. Heinze (2004) Knowing your enemies: seasonal dynamics of host-parasite recognition. Naturwissenschaften 91: 594-597.

21. T. Wenseleers, A.G. Hart, F.L.W. Ratnieks & J.J.G. Quezada-Euan (2004) Queen execution and caste conflict in the stingless bee Melipona beecheii. Ethology 110: 725-736.
Movie of a virgin Melipona beecheii queen being killed by workers (mpeg format)
Movie of a virgin Melipona beecheii queen being killed by workers (windows media format)

20. T. Wenseleers, A.G. Hart & F.L.W. Ratnieks (2004) When resistance is useless: policing and the evolution of reproductive acquiescence in insect societies. American Naturalist 164: 154-167.

19. T. Wenseleers, H. Helanterä, A.G. Hart & F.L.W. Ratnieks (2004) Worker reproduction and policing in insect societies. An ESS analysis. Journal of Evolutionary Biology 17: 1035-1047.

18. T. Wenseleers & F.L.W. Ratnieks (2004) Tragedy of the commons in Melipona bees. Proceedings of the Royal Society of London Series B-Biological Sciences 271: S310-S312. View talk on this (powerpoint)

17. T. Wenseleers (2004) The puzzle of cooperation. Trends in Ecology & Evolution 19: 409-410. (Review of Genetic and Cultural Evolution of Cooperation (P. Hammerstein, editor) and Foundations of Human Sociality (J. Henrich, R. Boyd, S. Bowles, C. Camerer, E. Fehr & H. Gintis, editors))

16. A.G. Hart, R.W. Bowtell, W. Kockenberger, T. Wenseleers & F.L.W. Ratnieks (2003) Magnetic resonance imaging in entomology: a critical review. Journal of Insect Science 3.5, 9 pages. Available online: www.insectscience.org/3.5 Accompanying files:
MRI sections of wasp abdomen
MRI sections of whole wasp
Reconstruction of wasp abdomen

15. A. Schotanus & T. Wenseleers (2003) Big Business: Bijenkolonies en oliemaatschappijen. Maandblad van de Vlaamse Imkerbond 89: 304-306. (in Dutch)

14. A. Schotanus & T. Wenseleers (2003) Conflict in het bijenvolk. Maandblad van de Vlaamse Imkerbond 89: 186-188. (in Dutch)

13. T. Wenseleers, F.L.W. Ratnieks & J. Billen (2003) Caste fate conflict in swarm-founding social Hymenoptera: an inclusive fitness analysis. Journal of Evolutionary Biology 16: 647-658. View talk on this (powerpoint)

12. S. Van Borm, T. Wenseleers, J. Billen & J.J. Boomsma (2003) Cloning and sequencing of wsp encoding gene fragments reveals a diversity of co-infecting Wolbachia strains in Acromyrmex leafcutter ants. Molecular Phylogenetics and Evolution 26: 102-109.

11. T. Wenseleers, J. Billen & A. Hefetz (2002) Territorial marking in the desert ant Cataglyphis niger: does it pay to play Bourgeois? Journal of Insect Behavior 15: 85-93.

10. T. Wenseleers, L. Sundström & J. Billen (2002) Deleterious Wolbachia in the ant Formica truncorum. Proceedings of the Royal Society of London Series B-Biological Sciences 269: 623-629.

9. K.R. Foster, T. Wenseleers & F.L.W. Ratnieks (2001) Spite: Hamilton's unproven theory. Annales Zoologici Fennici 38: 229-238. (Volume dedicated to the memory of William Donald Hamilton)

8. H. Moo-Valle, J.J.G. Quezada-Euán & T. Wenseleers (2001) The effect of food reserves on the production of sexual offspring in the stingless bee Melipona beecheii (Apidae, Meliponini). Insectes Sociaux 48: 398-403.

7. S. Van Borm, T. Wenseleers, J. Billen & J.J. Boomsma (2001) Wolbachia in leafcutter ants: a widespread symbiont that may induce male killing or incompatible matings. Journal of Evolutionary Biology 14: 805-814.

6. T. Wenseleers (2001) An introduction to the evolutionary modelling of conflict. In: "The Mathematical Modelling of Social Behaviour" (T. Wenseleers & F.L.W. Ratnieks, eds.), p. 21-53, EU INSECTS training workshop, Great Hucklow, UK, 14-18 Sept. 2001, 113 pages. Overheads Mathematica notebooks

5. K.R. Foster, F.L.W. Ratnieks & T. Wenseleers (2000) Spite in social insects. Trends in Ecology and Evolution 15: 469-470.

4. D. Grasso, T. Wenseleers, A. Mori, F. Le Moli & J. Billen (2000) Thelytokous worker reproduction and lack of Wolbachia infection in the harvesting ant Messor capitatus. Ethology, Ecology & Evolution 12: 309-314.

3. T. Wenseleers & J. Billen (2000) No evidence for Wolbachia-induced parthenogenesis in the social Hymenoptera. Journal of Evolutionary Biology 13: 277-280.

2. T. Wenseleers, F. Ito, S. Van Borm, R. Huybrechts, F. Volckaert & J. Billen (1998) Widespread occurrence of the micro-organism Wolbachia in ants. Proceedings of the Royal Society of London Series B-Biological Sciences 265: 1447-1452.

1. T. Wenseleers, E. Schoeters, J. Billen & R. Wehner (1998) Distribution and comparative morphology of the cloacal gland in ants (Hymenoptera: Formicidae). International Journal of Insect Morphology and Embryology 27: 121-128.

T. Wenseleers (2001) Conflict from Cell to Colony. Ph.D. thesis, University of Leuven, Belgium, 205 pp. Advisor: Prof. Dr. J. Billen.

B. Gobin, T. Wenseleers & J. Billen (1997) Microorganisms in the reproductive organs of the ant Gnamptogenys menadensis (Mayr) (Hymenoptera: Formicidae). In: "Regulation of reproduction and dominance interactions in the ponerine ant Gnamptogenys menadensis", Ph.D. thesis B. Gobin, University of Leuven, Belgium, pp. 57-60.

T. Wenseleers (1995) Exocrine glands as behavioural modulators in ants: the chemoecology of territorial marking in the desert ant Cataglyphis niger. Undergraduate thesis, University of Leuven, Belgium. Supervisors: Prof. Dr. J. Billen, Prof. A. Hefetz. Received a prize for best undergraduate thesis of the department

89. T. Wenseleers (2009) The evolution of conflict and cooperation in insect societies: towards greater realism in inclusive fitness models. First Meeting of the Central European Section of the IUSSI, Frauenchiemsee, Germany, 9-12 October 2009. (plenary)

88. D.A. Alves, V.L. Imperatriz-Fonseca, T. Francoy, P.S. Souza-Filho & T. Wenseleers (2009) Diploid male production in a small and isolated population of Brazilian stingless bee Melipona scutellaris. 41th Apimondia Congress, Montpellier, France, 15-20 September 2009.

87. T. Wenseleers (2009) Culturele evolutie als Darwinistisch proces. Of hoe methoden uit de biologie kunnen aangewend worden binnen de cultuurwetenschappen. Zomerschool "Evolutietheorie: verworvenheden en uitdagingen", KBIN, Brussel, 20 augustus 2009.

86. V.L. Imperatriz-Fonseca, D.A. Alves, P. Nogueira-Neto & T. Wenseleers (2009) Stingless bees as pollinators and ecosystem service providers in Brazilian agriculture. 10th International Congress of Ecology, Brisbane, Australia, 16-21 August 2009.

85. D.A. Alves, V.L. Imperatriz-Fonseca, T. Francoy, J. Billen & T. Wenseleers (2009) Intraspecific social parasitism by queens in the stingless bee Melipona scutellaris. 46th Annual Meeting of the Animal Behavior Society, Pirenpolis, Brazil, 22-26 June 2009.

84. T. Wenseleers (2009) De triomfen van de sociobiologie. College De Valk, K.U.Leuven, 16 June 2009. (lezingenreeks De wetenschap van evolutie)

83. T. Wenseleers (2009) Evolution and its Role in Science and Society. Institute for Advanced Study, São Paulo, Brazil, 18 March 2009. (invited talk) Available online as a streaming video.

82. T. Wenseleers (2009) Altruism in insect societies and beyond: voluntary or enforced? University of São Paulo, Brazil, 3 March 2009. (invited talk)

81. T. Wenseleers , D.A. Alves, T. Francoy, V.L. Imperatriz-Fonseca, P. Nogueira Neto & J. Billen (2009) Social parasitism in stingless bees. Royal Society Satellite Meeting on Social Insects, Sussex University, Brighton, UK, 20-21 January 2009.

80. W. Bonckaert & T. Wenseleers (2008) Worker policing in the two Vespine wasps Vespula vulgaris and Dolichovespula norwegica. Symposium "Entomology in Belgium 2008", K.B.I.N., Brussels, 5 December 2008.

79. U. Ernst , D. Cardoen, P. Verleyen, T. Wenseleers, D. de Graaf & L. Schoofs (2008) Worker policing in the honeybee Apis mellifera and other social insects. Symposium "Entomology in Belgium 2008", K.B.I.N., Brussels, 5 December 2008.

78. A. Vantaux , J. Billen & T. Wenseleers (2008) Cooperation and conflict in an ant-aphid mutualism: do ants sanction uncooperative aphid clones? Symposium "Entomology in Belgium 2008", K.B.I.N., Brussels, 5 December 2008.

77. T. Wenseleers , D.A. Alves, T. Francoy, V.L. Imperatriz-Fonseca, P. Nogueira Neto & J. Billen (2008) Intraspecific social parasitism by queens in stingless bees. Symposium "Entomology in Belgium 2008", K.B.I.N., Brussels, 5 December 2008.

76. T. Wenseleers (2008) Altruism in insect societies: voluntary or enforced? Biodiversity Research Centre, Université Catholique de Louvain, 13 November 2008. (invited talk)

75. T. Wenseleers (2008) Altruism in insect societies and beyond: voluntary or enforced? Universidad Nacional de Colombia, Medellin, Colombia, 26 September 2008. (invited talk)

74. T. Wenseleers (2008) Altruism in insect societies and beyond: voluntary or enforced? Universidad del Valle, Cali, Colombia, 24 September 2008. (invited talk)

73. T. Wenseleers (2008) Altruism in insect societies and beyond: voluntary or enforced? 14th EMPSEB meeting, Einsiedeln, Switzerland, 8-13 September 2008. (plenary)

72. U. Ernst , T. Wenseleers, P. Verleyen, D. Cardoen, D. de Graaf, L. Schoofs & F.L.W. Ratnieks (2008) Allocation of worker policing in the honeybee Apis mellifera, Proceedings of EurBee3, The 3rd European Conference of Apidology, Queen's University, Belfast, UK, 8-11 September 2008, p. 31.

71. W. Bonckaert & T. Wenseleers (2008) Worker policing in the two Vespine wasps Vespula vulgaris and Dolichovespula norwegica. 4th European Meeting of the IUSSI, La Roche en Ardenne, Belgium, 30 August-4 September 2008.

70. U. Ernst , T. Wenseleers, P. Verleyen, D. Cardoen, L. Schoofs & F.L.W. Ratnieks (2008) Worker policing in the honeybee – a job for a specialist? 4th European Meeting of the IUSSI, La Roche en Ardenne, Belgium, 30 August-4 September 2008.

69. T. Wenseleers (2008) Altruism in insect societies and beyond: voluntary or enforced? 4th European Meeting of the IUSSI, La Roche en Ardenne, Belgium, 30 August-4 September 2008. (plenary)

68. T. Wenseleers (2008) Altruism in insect societies and beyond: voluntary or enforced? 20th International Congress of Zoology, Paris, 26-29 August 2008. (plenary)

67. T. Wenseleers (2008) Altruism in insect societies and beyond: voluntary or enforced? International Congress of Entomology, Durban, South Africa, 6-12 July 2008.

66. T. Wenseleers (2008) Evolution and its role in science and society. Leuven Center for Bio-Science, Bio-Engineering and Bio-Technology (BioSCENTer) Launch Event, K.U.Leuven, 28 May 2008. (invited talk)

65. W. Bonckaert & T. Wenseleers (2007) Law and order in insect societies: the evolution of worker policing. Bee Lateral Symposium II, Leuven, Belgium, 17 December 2007.

64. T. Wenseleers (2007) Altruism in insect societies and beyond: voluntary or enforced? Université Paris-Nord, Paris, France, 12 October 2007. (invited talk)

63. T. Wenseleers (2007) Altruism in insect societies: voluntary or enforced? University of Caracas, Venezuela, 2 October 2007. (invited talk)

62. T. Wenseleers (2007) Altruism in insect societies and beyond: voluntary or enforced? 6th Meeting of the Bolivarian Section of the IUSSI, Higuerote, Venezuela, 25-27 September 2007. (plenary)

61. T. Wenseleers (2007) Altruism in insect societies: voluntary or enforced? University of Bern, Switzerland, 11 April 2007. (invited talk)

60. T. Wenseleers (2007). Coöperatief gedrag bij insecten: vrijwillig of gedwongen? Scientica symposium , K.U.Leuven, 20 March 2007. (invited talk)

59. T. Wenseleers (2007) Altruism in insect societies: voluntary or enforced? University of Copenhagen, Denmark, 15 March 2007. (invited talk)

58. T. Wenseleers (2007) Coöperatief gedrag bij insecten: vrijwillig of gedwongen? NecoV Wintersymposium, Nijmegen, The Netherlands, 18 January 2007. (plenary)

57. T. Wenseleers (2006) The social Hymenoptera: a model in the study of the evolution and genetics of social behaviour. Bee Lateral Symposium, Leuven, Belgium, 12 December 2006.

56. T. Wenseleers (2006) The origin of complex social behaviour: from insect societies to human cooperation and beyond. K.U.Leuven, 7 December 2006. Lecture given in the context of the award of the Prize of the Research Council (Prijs van de Onderzoeksraad).

55. T. Wenseleers (2006) Een wespennest van intriges: twisten en conflicten in insectenkolonies. PDL symposium, University of Leuven, 18 November 2006. (invited talk)

54. T. Wenseleers (2006) The effect of relatedness on the evolution of altruism in insect societies. 13th Benelux Congress of Zoology, Leuven, Belgium, 27-28 October 2006.

53. H. Helanterä, A. Tofilski, T. Wenseleers & F.L.W. Ratnieks (2006) Worker policing in the common wasp is not hygienic behaviour. 15th International Congress of the IUSSI, Washington DC, 30 July-4 August 2006.

52. T. Wenseleers (2006) Direct and indirect effects of kinship on conflict resolution in insect societies. 15th International Congress of the IUSSI, Washington DC, 30 July-4 August 2006.

51. T. Wenseleers (2006) The social insects' altruism: deliberate or enforced? University of Lausanne, Switzerland, 17 May 2006. (invited talk)

50. T. Wenseleers (2006) The social insects' altruism: deliberate or enforced? Spring Symposium, University of Helsinki, Finland, 8 March 2006. (plenary)

49. T. Wenseleers (2005) What can biology teach us about the way we are? Workshop Interdisciplinair Centrum voor Evolutie en Gedrag (ICEG), Leuven, Belgium, 5-6 September 2005.

48. T. Wenseleers (2005) How do insect societies resolve their conflicts? Institute for Theoretical Biology , Humboldt University Berlin, Germany, 13 July 2005. (invited talk)

47. T. Wenseleers (2005) From genes to societies: a biologist's view on conflict and cooperation. Institute for Advanced Study, Wissenschaftskolleg zu Berlin, 7 June 2005. (invited talk)

46. T. Wenseleers (2005) Conflict resolution in insect societies. Symposium "Conflict Resolution: Bringing Together Biologists and Social Scientists", Institute for Advanced Study, Wissenschaftskolleg zu Berlin, Germany, 4-7 May 2005. (plenary)

45. T. Wenseleers (2004) Are workers in highly eusocial insects altruistic or oppressed? University of Oulu, Finland, 15 September 2004. (invited talk)

44. T. Wenseleers (2004) Testing the importance of kinship and coercion in resolving conflicts in insect societies. Closing symposium of the EU Research Training Network "INSECTS", Helsingør, Denmark, 2-6 September 2004.

43. T. Wenseleers (2004) Conflict over female caste fate in insect societies. Université Pierre et Marie Curie , Paris, France, 23 April 2004. (invited talk)

42. T. Wenseleers (2004) The insect societies' police state. Dept. of Biology, K.U.Leuven, 20 February 2004.

41. T. Wenseleers (2004) Conflict over female caste fate in stingless bees. University of São Paulo, Brazil, 23 January 2004. (invited talk)

40. T. Wenseleers , A.G. Hart & F.L.W. Ratnieks (2003) Les abeilles trichent sur la determination de leur caste: la production de reines naines chez labeille sans dard Schwarziana quadripunctata. Meeting of the French Section of the IUSSI, Brussels, 1-3 September 2003.

39. F.L.W. Ratnieks & T. Wenseleers (2003). Conflict resolution by worker policing in insect societies. 9 th ESEB Congress, Leeds, UK, 19-23 August 2003. (plenary)

38. T. Wenseleers , A.G. Hart & F.L.W. Ratnieks (2003) On the evolution of acquiescence or why consent to social oppression. Midterm Review Meeting, EU Research Training Network "INSECTS", Laufen, Germany, 28-31 March 2003.

37. T. Wenseleers (2003) Wolbachia: microbial manipulator of insect reproduction. Chester College, UK, 21 March 2003. (invited talk)

36. T. Wenseleers (2002) Actual and potential tragedies: conflict over female caste fate in Apis and Melipona bees. University of Groningen, The Netherlands, 18 December 2002. (invited talk)

35. T. Wenseleers (2002) Actual and potential tragedies: conflicts over female caste fate in Apis and Melipona bees. Dept. of Biology, K.U.Leuven, 16 December 2002.

34. T. Wenseleers (2002) Actual and potential tragedies: conflict over female caste fate in stingless bees and honey bees. Winter meeting of the British section of the IUSSI, London, 6 December 2002.

33. T. Wenseleers (2002) Wolbachia as a source of intragenomic conflict in ant colonies. Minisymposium on Molecular Ecology, University of Helsinki, 7 November 2002. (plenary)

32. T. Wenseleers (2002) Conflict over female caste fate in Apis and Melipona bees. University of Sheffield, UK, 31 October 2002. (invited talk)

31. T. Wenseleers (2002) The anarchistic societies of the bees without sting, Central Association of Beekeepers 2002 Conference, Royal Leamington Spa, 13 October 2002. (plenary)

30. T. Wenseleers (2002) Conflict over caste fate in stingless bees. In : Anais do V Encontro Sobre Abelhas, V Encontro Sobre Abelhas, Ribeirão Preto, SP, Brazil, 4-7 September 2002, pp. 23-24.

29. T. Wenseleers & F.L.W. Ratnieks (2002) Conflict over caste fate in social insects: a tragedy of the commons examined. 14th International Congress of the IUSSI, Sapporo, Japan, 28 July – 3 August 2002.

28. T. Wenseleers (2002) Wolbachia as a source of intragenomic conflict in ant colonies. 14th International Congress of the IUSSI, Sapporo, Japan, 28 July – 3 August 2002.

27. A. Hart , T. Wenseleers, F.L.W. Ratnieks, R. Bowtell & W. Kockenberger (2002) Nuclear magnetic resonance imaging: an introduction. EU "INSECTS" Network Training Workshop "Morphological and histological techniques", Leuven, Belgium, 29-30 June 2002.

26. T. Wenseleeers (2002) Wolbachia : bacterial manipulator of insect reproduction. University of Sheffield, UK, 21 February 2002. (invited talk)

25. T. Wenseleers (2002) Wolbachia as a source of intragenomic conflict in ant colonies. EU "INSECTS" Network Training Workshop "Genetic Techniques", Oulu, Finland, 15-17 February 2002.

24. T. Wenseleers (2001) Conflict over caste fate in social insects: a tragedy of the commons examined. Rice University, Houston, USA, 3 December 2001.

23. T. Wenseleers , S. van Borm, J. Billen, L. Sundström, J.J. Boomsma & A. Buschinger (2001) Does Wolbachia manipulate ant reproduction? The current evidence reviewed. 2nd European Meeting of the IUSSI, Berlin, Germany, 25-29 September 2001.

22. T. Wenseleers (2001) Conflict from Cell to Colony, EU "INSECTS" Network Training Workshop "The Mathematical Modelling of Social Behaviour", Great Hucklow, UK, 14-18 September 2001. (plenary)

21. T. Wenseleers (2001) An introduction to the evolutionary modelling of conflict. In: The Mathematical Modelling of Social Behaviour , EU 'INSECTS' TMR network Training Workshop , Great Hucklow, UK, 14-18 September 2001, pp. 21-53.

20. T. Wenseleers (2000) Conflicts in cells and colonies : common principles and idiosyncratic differences. University of Lausanne, Switzerland, 4 December 2000. (invited talk)

19. T. Wenseleers (2000) Conflicts in cells and colonies : common principles and idiosyncratic differences. University of Sheffield, UK, 6 October 2000. (invited talk)

18. T. Wenseleers (2000) Conflicts in cells and colonies : common principles and idiosyncratic differences. University of Helsinki, Finland, 20 September 2000. (invited talk)

17. T. Wenseleers (2000) Conflicts in cells and colonies : common principles and idiosyncratic differences. 6th EMPSEB meeting, Vaalbeek, Belgium, 14-18 March 2000.

16. T. Wenseleers & J. Billen (2000) Conflict from cell to colony – Wolbachia in a social ant host. EU TMR workshop "Social evolution: an integrated study of the effects of kinship, communication, productivity and disease", Zürich, Switzerland, 21-24 January 2000.

15. T. Wenseleers & J. Billen (1999) Wolbachia in a social ant host: conflicts from cell to colony. EU TMR workshop "Social evolution: an integrated study of the effects of kinship, communication, productivity and disease", Castleton, Losehill Hall, UK, 23-27 July 1999.

14. E. van Walsum , T. Wenseleers & J. Billen (1999) Aggression in queenless worker groups: reproductive competition or social policing? EU TMR workshop "Social evolution: an integrated study of the effects of kinship, communication, productivity and disease", Castleton, Losehill Hall, UK, 23-27 July 1999.

13. T. Wenseleers & J. Billen (1998) Conflict from cell to colony : Wolbachia in a social ant host. 13th International Congress of the IUSSI, Adelaide, Australia, 29 December 1998-4 January 1999.

12. S. Van Borm , T. Wenseleers & J. Billen (1998) The selfish cytoplasmic micro-organism Wolbachia : widespread occurrence in ants, 5th Benelux Congress of Zoology, Ghent, Belgium, 6-7 November 1998.

11. T. Wenseleers , K.R. Foster, F.L.W. Ratnieks & J. Billen (1998) Spite: an important evolutionary phenomenon, 5th Benelux Congress of Zoology, Ghent, Belgium, 6-7 November 1998.

10. T. Wenseleers (1998) Conflict from cell to colony: the selfish microorganism Wolbachia in a social ant host. Belgian Royal Society of Zoology, K.B.I.N., Belgium, 7 October 1998. (invited talk)

9. T. Wenseleers (1998) Conflict from cell to colony - the selfish microorganism Wolbachia in a social ant host. University of Sheffield, UK, 27 May 1998. (invited talk)

8. T. Wenseleers & J. Billen (1998) Wolbachia in a social ant host : genomic conflicts and natural selection in a hierarchical system. Mathematical Biology of Pattern and Process Meeting, Bath, UK, 5-8 April 1998. Received a prize for the best lecture

7. T. Wenseleers & J. Billen (1998) Reproductive conflicts in social insects – the microorganism Wolbachia as a previously unrecognized party, 4th EMPSEB Meeting, Heraklion, Crete, 1-3 April 1998.

6. T. Wenseleers & J. Billen (1997) Reproductive conflict in social insects - the microorganism Wolbachia as a previously unrecognised party. 4th Benelux Congress of Zoology, Utrecht, The Netherlands, 14-15 November 1997.

5. T. Wenseleers & J. Billen (1997) Wolbachia and other selfish cytoplasmic elements in the aculeate Hymenoptera: some evolutionary implications. EU TMR workshop "Social evolution: an integrated study of the effects of kinship, communication, productivity and disease", Mols, Denmark, 6-9 June 1997.

4. T. Wenseleers , B. Gobin & J. Billen (1996) Sociogram: a generally applicable computer program for the visualisation and analysis of dominance interactions. 3rd Benelux Congress of Zoology, Namur, Belgium, 8-9 November 1996.

3. T. Wenseleers , B. Gobin & J. Billen (1996) A general approach to the visualisation and analysis of dominance interactions. 20th International Congress of Entomology, Florence, Italy, 25-31 August 1996. (poster)

2. T. Wenseleers , J. Billen & A. Hefetz (1995) Territorial behaviour in the ant Cataglyphis niger : does it pay to play Bourgeois? 2nd Benelux Congress of Zoology, Leiden, The Netherlands, 16-17 November 1995.

1. T. Wenseleers , J. Billen & A. Hefetz (1995) Territorial behaviour in the ant Cataglyphis niger : does it pay to play Bourgeois? Meeting of the German section of the IUSSI, Utrecht, The Netherlands, 31 August 1995.

Worker bees cheat on queen behind her back so they can have an easier life. The Daily Mail, Indopia September 2009.

Worker bees cheat on queen. The Asian Age September 2009.

Sex Wars: Class Struggle In Bees. ScientificBlogging September 2009.

Die Gnade der heimlichen Geburt. Süddeutsche Zeitung September 2009.

Klassenkampf hält Bienen jung. ScienceTicker September 2009.

Lotta di classe nell'alveare. Le Scienze September 2009.

New research discovers worker bees in 'reproductive class war' with queen. EurekAlert!, ScienceCentricNews, AlphaGalileo, E-ScienceNews.Com, FirstScience, PhysOrg, RedOrbit, ScienceDaily, ScienceBlog September 2009.

Worker bees engage in 'reproductive class war' with queen. Yahoo News, Top News, South Asian News, One India, NetIndia 123, Thaindian News, Sindh Today, LittleAbout September 2009.

God Planned Bees to Lie and Cheat. The Detroit News September 2009.

Some worker bees 'reproduce for themselves'. The Telegraph, Big News Day September 2009.

S. Voormolen (2008) De vriendelijke mier. Sociobioloog Edward Wilson keert zich tegen verwantenselectie. NRC Handelsblad, January 2008. Commentary on recent articles by Edward Wilson in Social Research, PNAS, Quarterly Review of Biology and Bioscience, see also my own published critique in Trends in Ecology & Evolution.

Willy van Strien (2008) Het penisduel en ander strategisch gedrag van dieren. Uitgeverij KNNV, 240 pp, 19.95 euro. Excellent book written by freelance science writer Willy van Strien and discussing some of my research. Order a copy

P. Tyson (2007) Being queen. NOVA, October 2007.

Ong thợ bị tẩy não. Think.Com.Vn August 6 2007. (in Vietnamese)

Những công nhân bị tẩy não. Diễn Đàn Doanh Nghiệp, August 16 2007. (in Vietnamese)

J. Turney (2007) Helping out: altruism and cooperation. Wellcome Science February 2007.

CD of Vaughan Williams: Symphonies Nos. 4-6, including the piece "The Wasps (Overture)" (features one of my lovely wasp photos on the cover).
(note the opinion of one reviewer: "I have never liked "The Wasps" very much, but it is given a fine performance and you get a garish photo of a lot of wasps on the front of the CD. Don't hesitate. This is a great CD."
Want to buy it? Order a copy at Amazon!

D. Draulans (2007) De knuppel in de bijenkorf. Knack februari 2007.

M. Crok (2007) Terreur in de bijenkorf. Natuur, Wetenschap & Techniek februari 2007.

M. Roldan (2007) Altruismo o coaccion? Evolucion January 2007.

W. van Strien (2007) Acht verklaringen voor altruisme. Bionieuws 20 januari 2007.

Bijen leven in politiestaat. Quest Magazine januari 2007.

Insect Research: Some honeybees are enforcers of worker bee's eggs. Life Science Weekly, 26 December 2006.

Honeybees make ruthless relatives. MSNBC 12 December 2006.

Why ants, bees and wasps eat their siblings' eggs. FOX News 11 December 2006.

C. Binns (2006) Why bees eat their kin. Live Science 11 December 2006.

Vervenne speelt Sinterklaas. Veto 11 december 2006.

Vier universitaire onderzoekers in de prijzen. Het Nieuwsblad 8 december 2006.

Onderzoeksprijzen universiteit. Het Laatste Nieuws 8 december 2006.

Kin selection theory upheld. Press Display, 9 December 2006.

Why do some queen bees eat their worker bee's eggs? Science Daily, Biology News, Innovations Report, Biology Daily.com, First Science, GlobalSpec, PhysOrg.com, EurekAlert, SpiritIndia.com 6 December 2006.

Why do some queen bees eat their worker bee's eggs? Kisti (in Korean) 11 December 2006.

D.C. Queller (2006) To work or not to work. Nature 444: 42-43.

J. Duffy (2006) Anti-social bee-behaviour. BBC News Magazine, 3 November 2006.

B. Harder (2006) No reproductive rights in insect "police states", study finds. National Geographic News, Digg.com, Broz News, Cyanova Life Science News 1 November 2006.

Egg police crack down on broody bees. New Scientist 2576: 16 (1 November).

N. Fleming (2006) Why bees abide by the law. The Daily Telegraph, NewsVine 2 November 2006.

Scientists study insect "egg police". United Press International, Science Daily, Political Gateway, Ama Techtel, Digis, News Vantage, Monsters and Critics, Rootly.com, DailyIndia.com, WebIndia123.com, PhysOrg.com, Physics News 8 November 2006.

N. Anthis (2006) Social insects: More KGB than brady bunch. The Scientific Activist 16 November 2006.

S. Anitei (2006) Dictatorship and not voluntary altruism characterizes insect colonies. Worker females are forbidden to reproduce. Softpedia 13 November 2006.

S. Milius (2006) Tough policing deters cheating in insects. Science News 25 November 2006.

K. De Rijck (2006) Sociale sancties werken. De Standaard 2 november 2006.

J. Poot (2006) Jonge onderzoekers in de prijzen. "Insectengemeenschappen eerder politiestaat dan democratie". Campuskrant 22 november 2006.

Altruïsme onder dwang. Teleac, 2 november 2006.

Altruïsme werksterbijen is kwestie van dwang. De Volkskrant, 1 november 2006.

R. Bonalume Neto (2006) Abelhas patrulham ninho para controlar operárias. Folha de São Paulo 7 November 2006.

R. Bonalume Neto (2006) Repressão evita ato criminoso, diz cientista. Folha de São Paulo 7 November 2006.

Ramiro Velásquez Gómez (2006) Insectos decretan pena de muerte. El Colombiano (Colombia) 8 November 2006.

G. Pesthy (2006) „Rendőrállam” a társas rovaroknál (in Hungarian). National Geographic Magazine (Hungary), 3 November 2006.

Kaptárrendőrség (in Hungarian). Duna Television, Metro, KarpatInfo, RexInfo (Hungary), 10 November 2006.

"Rendőrök" pusztítják a dolgozók petéit (in Hungarian). Hirado, MTV Magazin (Hungary), 10 November 2006.

Social insect altruism is maintained by policing methods (in Russian). Elementy (Russia) 8 November 2006.

No reproductive rights in insect "police states", study finds (in Korean). Kisti (Korea) 6 November 2006.

Tvång och våld styr de arbetande bina (in Swedish). Arbetarbladet (Sweden) 14 November 2006.

Social kontroll styr arbetarbin (in Swedish). Metro (Sweden) 13 November 2006.

Samhällen styrs med våld och tvång (in Swedish). Helsingborgs Dagblad (Sweden) 15 November 2006.

Bích Vân (2006) Kiểm soát nghiêm nhặt đến độ nào trong xã hội côn trùng? (in Vietnamese) Khoa Hoc (Vietnam) 16 November 2006.

Presja społeczna odbiera osom macierzyństwo ("Social pressure prevents wasps from having offspring") (in Polish). Wiadomosci, Student News (Poland) 3 November 2006.

Pondre, une attitude punie chez les ouvrières insectes vivant en société. Le Monde, Romandie, Orange.fr, AOL.fr (France), La Presse, Le Nouvelliste, Cyberpresse (Canada), La Liberté, Bluewin, Edicom, TSR, SwissInfo (Switzerland), 1 November 2006.

Les ouvrières s’interdisent de pondre. L'Agefi (Switzerland) 3 November 2006.

Diktatur im Bienenstock. n-tv.de 1 November 2006.

Arbeiter im Insektenstaat werden kontrolliert. N24, Sat. 1 News, Netzeitung.de, 2 November 2006.

Soziale Kontrolle erzwingt Selbstlosigkeit bei Bienen. Donaukurier, Handelsblatt, Fränkischer Tag, Leipziger Volkszeitung, RNZ Online, Rhein-Neckar Zeitung, Schwäbische Zeitung, 3Sat, Mainpost.de, PNP Online, Sol.de, Main-Rheiner, Neue Osnabruecker Zeitung, Suedkurier, Neue Westfälische, Walsroder Zeitung, Mindelheimer Zeitung, Sueddeutsche Zeitung, news.yahoo.de, Oekokraft.de 1 November 2006.

Bienen erzwingen soziales Verhalten. In Insektenstaaten zahlt sich Egoismus nicht aus. Berliner Zeitung 2 November 2006.

Strenge soziale Kontrolle bei Bienen und Wespen. Bremer Nachrichten 6 November 2006.

Reino Unido: Insectos deixam de pôr ovos por coerção, diz estudo britânico. Angola Press, Yahoo Brasil Noticias, Terra Noticias, Último Segundo (Brazil), 2 November 2006.

Insectos dejan de poner huevos por miedo a ser asesinadas. La Crónica (Mexico), 2 November 2006.

Policía de Estado. El Diario de Chihuahua (Mexico) 26 November 2006.

Insectos dejan de poner huevos por coerción. La Patria (Bolivia), 12 November 2006.

Científicos prueban que algunos insectos dejan de poner huevos por coerción. El Mundo (Spain) 2 November 2006.

Insectos reciben órdenes. TVN (Chile) 2 November 2006.

D. Brown (2006) Genoma de la abeja puede revelar nuestra evolución social. El Mercurio (Chile), 3 November 2006.

Unos científicos afirman que algunos insectos dejan de poner huevos al ser vigilados. El Diario de Mallorca, 2 November 2006.

R.J. Lopes (2006) Abelhas têm "polícia" contra ovos não-autorizados, dis estudo. G1, O Popular (Brazil), 1 November 2006.

Algunos insectos dejan de poner huevos por coerción. BVS (Cuba) 4 November 2006.

Control social hace que sólo la abeja reina se reproduzca. La Tercera (Chile) 1 November 2006.

Westliche Honigbiene. Entry in the Wikipedia encyclopedia referring to our recent Nature paper. November 2006.

Amazonenameise. Entry in the Wikipedia encyclopedia referring to my recent paper with Patrizia D'Ettorre. October 2006.

Spite. Entry in the Wikipedia encyclopedia referring to my recent paper with Kevin Foster and Francis Ratnieks. October 2006.

Group selection. Entry in the Wikipedia encyclopedia referring to my recent paper with Kevin Foster and Francis Ratnieks. March 2006.

Hoornaars gaan mountainbikers te lijf. Gazet van Antwerpen, 26 september 2006.

En de prijs gaat naar... Science@Leuven jaargang 5 nummer 13, sept. 2006, p. 19. (in verband met uitrijking Prijs van de Onderzoeksraad)

PDL minisymposium, aankondiging lezing "Een wespennest van intriges: twisten en conflicten in insectenkolonies" (18 nov. 2006, Groot Auditorium Zoologisch Instituut), Science@Leuven jaargang 5 nummer 13, sept. 2006.

R. Dunn (2006) The Social Life - What humans can learn from social insects. Seed Magazine, 31 August 2006.

A. Chana (2006) Pretenders to the throne. Cosmos Magazine, 30 August 2006.

Gene meanies let themselves go...on and on. New Zealand Herald, 8 April 2006.

R. Highfield (2006) Ruthless DNA. The Telegraph (Calcutta, India), 3 April 2006.

R. Highfield (2006) Loveable rogue, or selfish killer? The Daily Telegraph 21 March 2006, p. 27. (note me being mentioned on the same page as Richard Dawkins and Sir John Krebs :-) )

M. Price (2006) The controversy of group selection theory. The Science Creative Quarterly, March 2006.

J.E. Niven (2005) A Cinderella story. Journal of Experimental Biology 208: 5-6.

T.J. Daniels, H. Winet, A.A. Smith, et al. (2005) Why we invoke Darwin. The Scientist 19: 8-9.

Bug cops. The Wilson Quarterly, Summer 2005. June 22 2005.

Limburgse wespennesten in 2004 verdubbeld. De Standaard, May 27 2005, p. 23.

Limburgse wespenwereld uit de doeken gedaan in boekje. Het Belang van Limburg, May 26 2005, p. 18.

Cops with six legs. The Tribune, Chandigarh, India 8 April 2005.

Y. Martínez (2005) Las comunidades de insectos generan sus propios estados policiales. La coerción favorece la cooperación entre las abejas y las hormigas. Tendencias Científicas 28 March 2005.

L. Singh (2005) A bug's life - Orwell style. Imprint 25 March 2005.

S. Milius (2005) Cover story: Cops with six legs. Law and order among insects. Science News 167: 184-186.

Repeated Creations of Insect Police Systems. 12 March 2005. Reasons to Believe, 12 March 2005. (note how our study is abused to support creationism!)

M. Rauner (2005) Korruption im Bienenstock. Zeitwissen 2/2005.

Bienen leben im Polizeistaat. www.hymenoptera.de, 30 January 2005.

D. Whipple (2005) Blue Planet: The 'Age of Insects'. The Washington Times 26 Jan. 2005.

L. Spinney (2005) Anarchy discovered in the honeybee hive. New Scientist 15 Jan. 2005.

R. B. Neto (2005) Sexo, mentiras e politicagem. Jornal da Ciencia 17 Jan. 2005 and Folha de Sao Paulo 16 Jan. 2005.

Bienenkorb als Polizeistaat - Parallelen zwischen Menschen und Tieren. Deutschlandfunk 7 Jan. 2005.

K. Seefelt (2005) Demokratie, Herrschaft des Volkes? Nix da, Rübe ab! Telepolis 7 Jan. 2005.

D. Draulans (2005) Bijenpolitie. Knack Magazine 12 Jan 2005.

S. Blackman (2004) Spite: evolution finally gets nasty. The Scientist 18: 14.

N.W. Atckinson (2004) Too many princesses. Natural History Magazine Dec. 2004/Jan. 2005, p. 16.

Mierenkolonie van 100 kilometer teistert Australie. Het Laatste Nieuws, Aug. 13 2004, p. 19.

D. Draulans (2004) Ik wil koningin worden. Knack Magazine 21 April 2004.

C. Pfaff (2004) Familie oder Karriere? Auch Bienen müssen sich entscheiden. Wissenschaft.de 3 March 2004.

Berufswahl bei Bienen - Was will ich werden - Königin oder Arbeiterin? Tropenwaldznetwerk-Brasilien 2 March 2004.

M. Dawes (2003) Watch out for the Sting! The Sheffield Star 29 May, p. 14.

D. Draulans (2001) Wolbachia en conflicten in mierenkolonies, Knack Magazine 31, p. 91.

G0G41A    Theoretical modelling in biology

• Lesson 1: Introduction: why model, modelling approaches. References Guide to Mathematica Exercises with model solutions
• Lesson 2: Difference and differential equation models with applications in population ecology and evolution. Worked examples Exercises with model solutions
• Lesson 3: Exact genetic models and the analysis of class structured populations. Worked examples Exercises with model solutions
• Lesson 4: Optimisation and inclusive fitness models. Worked examples Exercises with model solutions
• Lesson 5: Numerical models. I. (Joost Vanoverbeke) Simile models Stochastic models in Mathematica
• Lesson 6: Numerical models. II. (Joost Vanoverbeke) Simile models
• Lesson 7: Numerical models. III. (Joost Vanoverbeke) Simile models

G9X21A    Ethology and behavioural ecology (Part II)

• Lesson 1: The Evolution of Communication (Chapter 9 in Alcock).
• Lesson 2: The Evolution of Reproductive Behaviour (Chapter 10 in Alcock).
• Lesson 3: The Evolution of Mating Systems (Chapter 11 in Alcock).
• Lesson 4: The Evolution of Parental Care (Chapter 12 in Alcock).
• Lesson 5: The Evolution of Social Behaviour (Chapter 13 in Alcock).
• Lesson 6: The Evolution of Human Behaviour (Chapter 14 in Alcock).

G0F87A    Data Analysis Applied to Biological Research

• Chapter 11: Multiway ANOVA.
• Chapter 12: Complex ANOVA designs.
• Chapter 15: Nonparametric statistics.
• Chapter 16: Power analysis.
• Exercises Complex ANOVA designs. exercises results datasets Excel formulae Sequential Bonferroni Benjamini-Hochberg
• Demonstration bootstrap analysis & permutation tests in Maple and Matlab. handout Maple file Matlab file (1) Matlab file (2)
• Exercises Nonparametric statistics and power analysis. exercises results datasets

G0I29a    Advanced Ecology: Field Course

• The behavioural ecology of seabirds.

G0G30A   Evolutietheorieën

• Darwiniaanse benaderingen van culturele evolutie: methoden en toepassingen. (guest lecture)

The evolutionary and physiological basis of altruism. Guest lecture in course "Comparative Physiology", University of Ghent, Belgium.

EvoGen workgroup

• How to control for phylogenetic non-independence in biological analyses: an update on the comparative method. June 2006.

Modern research methods in biology

• Determining kinship relationships using genetic markers, March 20 2006.

Selected topics in ethology (in Dutch)

• The evolution of altruism and highly eusocial behaviour. c
• The calculation of relatedness. handout
• The evolution of social conflicts. handout
• The evolution of human behaviour. handout
• Excercises and problems (to be known for exam) handout

The Social Wasps of Britain. Pamphlet. Page 1 Page 2

Bumblebees of the Sheffield Area. Pamphlet.

Wolbachia: microbal insect symbionts and manipulators of reproduction. Chester College.

Actual and potential tragedies: conflict over female caste fate in Apis and Melipona bees.

Wolbachia : microbial manipulator of insect reproduction. Guest lecture in the Selfishness & Altruism course.

An introduction to the evolutionary modelling of conflict, EU 'INSECTS' TMR network Training Workshop "The Mathematical Modelling of Social Behaviour", Great Hucklow, UK, 2001. Overheads Mathematica notebooks

The evolution of conflict and cooperation. Population biology and population genetics seminar series, University of Leuven, Belgium.

Werkcolleges statistische gegevensverwerking. Statistics manual, University of Leuven, Belgium, 121 pages. (in Dutch)